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A thorough analysis of the effect of the linker size Supplementary Note 5 showed that the shell thickness may appear non-determinable if the estimation of the linker size is incorrect. Kaufman HE.

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Virology42—53 This highlights that the use Hsv dating Cambridge primary labelling allows the measurement of smaller thickness that secondary labelling cannot reliably determine. The analysis of the capsid spatial offset is described in details in Supplementary Note 6.

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Resistance mutations identified in the UL5 helicase or UL52 primase genes were validated by marker transfer. We note that for VP16 and pUL37, the use of secondary labelling did not allow the estimation of the thickness parameter. For more complex structures, the alignment may require more complex algorithms, such as those reported for the alignment of nuclear pore complexes and discussion.